Only papars written in English are presented here.


Hikosaka K (2005a) Leaf canopy as a dynamic system: ecophysiology and optimality in leaf turnover. Annals of Botany, 95: 521-533.
Hikosaka K (2005b) Nitrogen partitioning in the photosynthetic apparatus of Plantago asiatica leaves grown at different temperature and light conditions: similarities and differences between temperature and light acclimation. Plant and Cell Physiology, 46: 1283-1290.
Hikosaka K, Takashima T, Kabeya D, Hirose T, Kamata N (2005a) Biomass allocation and leaf chemical defense in defoliated seedlings of Quercus serrata with respect to carbon-nitrogen balance. Annals of Botany, 95: 1025-1032.
Hikosaka K, Onoda Y, Kinugasa K, Anten NPR, Nagashima H, Hirose T (2005b) Plant responses to elevated CO2 concentration at different scales: leaf, whole plant, canopy, and population. Ecological Research, 20: 243-253.
Hirose T (2005) Development of the Monsi and Saeki theory on canopy structure and function. Annals of Botany 95: 483-494.
Kinugasa T, Hikosaka K, Hirose T (2005) Respiration and reproductive effort in Xanthium canadense. Annals of Botany, 96: 81-89.
Makino, T. T. and Sakai, S. (2005) Does interaction between bumblebees (Bombus ignitus) reduce their foraging area?: bee-removal experiments in a net cage. Behavioral Ecology and Sociobiolgy 57:617-622.
Muller O, Hikosaka K, Hirose T (2005) Seasonal changes in light and temperature affect the balance between light utilisation and light harvesting components of photosynthesis in an evergreen understorey. Oecologia, 143: 501-508.
Oguchi R, Hikosaka K, Hirose T (2005) Leaf anatomy as a contraint for photosynthetic acclimation: differential responses in leaf anatomy to increasing growth irradiance among three deciduous trees.Plant, Cell and Environment, 28: 916-927.
Oikawa S, Hikosaka K, Hirose T (2005) Dynamics of leaf area in a canopy of an annual herb, Xanthium canadense. Oecologia, 143: 517-526.
Onoda Y, Hikosaka K, Hirose T (2005a) Seasonal change in the balance between capacities of RuBP carboxylation and RuBP regeneration affects CO2 response of photosynthesis in Polygonum cuspidatum. Journal of Experimental Botany, 56: 755-763.
Sakai, S. and Harada, Y. (2005) Production of offspring using current income and reserves: size-number trade-off and optimal offspring size. Journal of theoretical Biolgy 233:65-73.
Sakai, S. and Sakai, A. (2005) Nature of size-number trade-off: test of the terminal-stream-limitation model for seed production of Cardiocrium Cordatum. Oikos 108:105-114.
Wright IJ, Reich PB, Cornelissen JHC, Falster DS, Garnier E, Hikosaka K, Lamont BB, Lee W, Oleksyn J, Osada N, Poorter H, Villar R, Warton DI, Westoby M (2005a) Assessing the generality of global leaf trait relationships. New Phytologist, 166: 485-496.
Wright IJ, Cornelissen JHC, Falster DS, Groom PK, Hikosaka K, Lee W, Lusk CH, Niinemets ?, Oleksyn J, Osada N, Poorter H, Reich PB, Warton DI, Westoby M (2005b) Modulation of leaf economic traits and trait relationships by climate. Global Ecology and Biogeography, 14: 411-421.
Yasumura Y, Onoda Y, Hikosaka K, Hirose T (2005) Nitrogen resorption from leaves under different growth irradiance in three deciduous woody species. Plant Ecology, 178: 29-37.



Anten NPR, Hirose T, Onoda Y, Kinugasa T, Kim HY, Okada M, Kobayashi K (2004) Elevated CO2 and nitrogen availability have interactive effects on canopy carbon gain in rice. New Phytologist, 161: 459-472
Hikosaka K, Kato MC, Hirose T (2004) Photosynthetic rates and partitioning of absorbed light energy in photoinhibited leaves. Physiologia Plantarum, 121: 699-708.
Iwaizumi, M. & Sakai, S. (2004) Variation in flower biomass among nearby populations in Impatiens textori (Balsaminaceae): effects of population plant densities. Canadian Journal of Botany 82:563-572.
Makino, T. T. & Sakai, S. (2004) Findings on spatial foraging patterns of bumblebees (Bombus ignitus) from a bee-tracking experiment in a net cage.Ý Behavioral Ecology and Sociobiology 56:155-163.
Oikawa S, Hikosaka K, Hirose T, Hori Y, Shiyomi M, Takahashi S (2004) Cost-benefit relationships in leaves emerging at different times in a deciduous fern Pteridium aquilinum. Canadian Journal of Botany, 82: 521-527.
Onoda Y, Hikosaka K, Hirose T (2004) Nitrogen allocation to cell walls decreases photosynthetic nitrogen-use efficiency. Functional Ecology, 18: 419-425
Sakai, S. and Harada, Y. (2004) Size-number trade-off and optimal offspring size for offspring produced sequentially using a fixed amount of reserves. Journal of theoretical Biology 226: 253-264.
Takashima T, Hikosaka K, Hirose T (2004) Photosynthesis or persistence: nitrogen allocation in leaves of evergreen and deciduous Quercus species. Plant, Cell and Environment, 27: 1047-1054.
Wright IJ, Reich PB, Westoby M, Ackerly DD, Baruch Z, Bongers F, Cavendar-Bares J, Chapin T, Cornelissen JHC, Diemer M, Flexas J, Garnier E, Groom PK, Gulias J, Hikosaka K, Lamont BB, Lee T, Lee B, Lusk C, Midgley JJ, Navas M-L, Niinemets , Oleksyn J, Osada N, Poorter H, Poot P, Prior L, Pyankov VI, Roumet C, Thomas SC, Tjoelker MG, Veneklaas EJ, Villar R (2004) The worldwide leaf economics spectrum. Nature, 428: 821-827.




Hikosaka K (2003) A model of dynamics of leaves and nitrogen in a plant canopy: An integration of canopy photosynthesis, leaf life span, and nitrogen use efficiency. American Naturalist, 162: 149-164.
Hikosaka K, Yamano T, Nagashima H, Hirose T (2003) Light-acquisition and use of individuals as influenced by elevated CO2 in even-aged monospecific stands of Chenopodium album. Functional Ecology, 17: 786-795.
Ishizaki S, Hikosaka K, Hirose T (2003) The increase in leaf mass per area benefits plant growth at elevated CO2 concentration. Annals of Botany, 91: 905-914.
Kabeya, D. and Sakai, S. (2003) The role of roots and cotyledons as storage organs in early stages of establishment in Quercus crispula: a quantitative analysis of the nonstructural carbohydrate in cotyledons and roots. Annals of Botany 92:537-545.
Kabeya, D., Sakai, A., Matsui, K., & Sakai, S. (2003) Resprouting ability of Quercus crispula seedlings depends on the vegetation cover of their microhabitats. Journal of Plant Research 116: 207-216.
Kato MC, Hikosaka K, Hirotsu N, Makino A, Hirose T (2003) The excess light energy that is neither utilized in photosynthesis nor dissipated by photoprotective mechanisms determines the rate of photoinactivation in photosystem II. Plant and Cell Physiology, 44: 318-325.
Kinugasa T, Hikosaka K, Hirose T (2003) Reproductive allocation of an annual, Xanthium canadense, at an elevated carbon dioxide concentration. Oecologia, 137: 1-9.
Morinaga, S.-I., Tsuji, K., and Sakai, S. (2003) Consequences of differences in flowering date in seed production in Heloniopsis orientalis (Liliaceae). American Journal of Botany 90: 1153-1158.
Oguchi R, Hikosaka K, Hirose T (2003) Does the photosynthetic light-acclimation need change in leaf anatomy? Plant, Cell and Environment, 26: 505-512.
Nagashima H, Yamano T, Hikosaka K, Hirose T (2003) Effects of elevated CO2 on the size structure in even-aged monospecific stands of Chenopodium album. Global Change Biology, 9: 619-629.
Sakai, A., Matsui, K. , Kabeya, D. & Sakai, S. (2003) Altitudinal variation in lifetime growth trajectory and reproductive schedule of a sub-alpine conifer Abies mariesii. Evolutionary Ecology Research 5: 671-689.
Sakai, A. & Sakai, S. (2003) Size-dependent ESS sex allocation in wind-pollinated cosexual plants: fecundity vs. stature effects. Journal of theoretical Biology 222: 283-295.
Tsonev TD, Hikosaka K (2003) Contribution of photosynthetic electron transport, heat dissipation, and recovery of photoinactivated photosystem II to photoprotection at different temperatures in Chenopodium album leaves. Plant and Cell Physiology, 44: 828-835.
Ushimaru, A., Itagaki, T. & Ishii, H.S. (2003) Variation in floral organ size is dependent on function: a test with Commelina communis, an andromonoecious species. Evolutionary Ecology Research 5: 615-622
Ushimaru, A., Itagaki, T. & Ishii, H.S. (2003) Floral correlations in an andromonoecious, Commelina communis (Commelinaceae). Plant Species Biology 18: 103-106



Hikosaka, K., Nagamatsu, D., Ishii, H.S., Hirose, T. (2002) Photosynthesis-nitrogen relationships in species at different altitudes of Mount Kinabalu, Malaysia. Ecological Research 17: 305-313.
Ishii, H. S. & Sakai, S. (2002) Temporal variation in floral display size and individual floral sex allocation in racemes of Narthecium asiaticum (Liliaceae). American Journal of Botany 89:441-446.
Kato, M.C., Hikosaka, K., & Hirose, T. (2002) Photoinactivation and recovery of photosystem II of Chenopodium album leaves grown under different irradiance and nitrogen availability. Functional Plant Biology (formerly Australian Journal of Plant Physiology) 29: 787-795.
Kato, M.C., Hikosaka, K., & Hirose, T. (2002) Leaf discs floated on water are different from intact leaves in photosynthesis and photoinhibition. Photosynthesis Research 72: 65-70.
Kimura, T., Ishii, H. S. & Sakai, S. (2002) Selfed-seed production depending on individual size and flowering sequence in Iris gracilipes (Iridaceae). Canadian Journal of Botany 80:1096-1102.
Nagamatsu, D., Seiwa, K. & Sakai, A. (2002) Seedling establishment of deciduous trees in various topographic positions. Journal of Vegetation Science13:35-44.
Werger, M.J.A., Hirose, T., During, H.J., Heil, G.W., Hikosaka, K., Ito, T., Nachinshonhor, Shibazaki, K., Nagamatsu, D., Takatsuki, S., & Anten, N.P.R. (2002) Changes in photon flux partitioning among species and species composition in grassland after cessation of grazing. Journal of Vegetation Science 13: 615-626.
Yasumura, Y., Hikosaka, K., Matsui, K., & Hirose, T. (2002) Leaf-level nitrogen use efficiency of species in deciduous forests. Functional Ecology 16: 826-834.


Anten, N.P.R. & Hirose, T. (2001) Limitations on photosynthesis of competing individuals in stands and consequences for canopy structure. Oecologia 129: 186-196
Hikosaka, K. & Hirose, T. (2001) Nitrogen uptake and use by competing individuals in a Xanthium canadense stand. Oecologia 126: 174-181.
Hikosaka, K., Nagashima, H., Harada, Y. & Hirose, T. (2001) A simple formulation of interaction between individuals competing for light in a monospecific stand. Functional Ecology 15: 642-646.
Ishii, H. S. & Sakai, S. (2001) Implications of geitonogamous pollination for floral longevity in Iris gracilipes (Iriddaceae). Functional Ecology 15:633-641.
Ishii, H. S. & Sakai, S. (2001) Effects of floral display size and floral positions on individual floral longevities in racemes of Narthecium asiaticum (Liliaceae). Functional Ecology 15:396-405
Ohashi, K. & Yahara, T. (2001) Behavioral responses of pollinators to variation in floral display size and their influences on the evolution of floral traits. In (eds by Chittaka L, Thompson JD) Cognitive ecology of pollination. pp. 274-296. Cambridge University Press.
Sakai, S. and Harada, Y. (2001) Why do large mothers produce large offspring?: theory and a test. American Naturalist 157:348-359.
Sakai, S. and Harada, Y. (2001) Sink-limitation and the size-number trade-off of organs: production of organs using a fixed amount of reserves. Evolution 55:467-476.



Anten, N.P.R., Hikosaka, K. & Hirose, T. (2000) Nitrogen utilisation and the photosynthetic system. In: eds. B Marshall and J Roberts, Leaf development and canopy growth. pp. 171-203, Sheffield Acedemic Press, Sheffield.
Hikosaka, K. & Hirose, T. (2000) Photosynthetic nitrogen use efficiency in species coexisting in a warm-temperate evergreen forest. Tree Physiology, 20: 1249-1254.
Ishii, H. S. & Sakai, S. (2000) Optimal timing of flower abortion; experimental study on Erythronium japoncum (Liliaceae). Functional Ecology14:122-128.
Ohashi, K. & Yahara, T. (2000) Effects of flower profuction and predispersal seed predation on reproduction in Cirsium purpuratum. Canadian Journal of Botany 78: 230-236.
Sakai, S. (2000) Biased sex allocation in hermaphroditic plants. Journal of Plant Research 113: 335-342.
Sakai, S. (2000) Size-number trade-off and allocation to flower production in animal-pollinated plants. Plant Species Biology 15: 261-268.



Anten, N.P.R. & Hirose, T. (1999) Intraspecific differences in above-ground growth patterns result in spatial and temporal partitioning of light among species in a tall-grass meadow. Journal of Ecology, 87: 583-597.
Hikosaka, K., Sudoh, S. & Hirose, T. (1999) Light acquisition and use of individuals competing in a dense satnd of an annual herb, Xanthium canadense. Oecologia, 118: 388-396.
Hikosaka, K., Murakami, A. & Hirose, T. (1999) Balancing carboxylation and regenaration of ribulose-1,5-bisphosphate in leaf photosynthesis in temperature acclimation of an evergreen tree, Quercus myrsinaefolia. Plant, Cell & Environment, 22: 841-849.
Nagashima, H. (1999) The processes of height-rank determination among individuals and neighbourhood effects in Chenopodium album L. stands. Annals of Botany, 83: 501-507.
Ohashi, K. & Yahara, T. (1999) How long to stay on, and how often to visit a flowering plant? - a model for foraging strategy when floral displays vary in size. OIKOS, 86: 386-392.
Sakai, S. (1999) Female-biased sexual allocation in cosexual plants: result of sink-limited growth of fruits. Evolutionary Ecology Research, 1: 953-957.
Sakai, S. & Ishii, H. S. (1999) Why be completely outcrossing? evolutionarily stable outcrossing strategies in an environment where outcross pollen availability is unpredictable. Evolutionary Ecology Research, 1: 211-222.


Anten, N.P.R. & Hirose, T. (1998) Biomass allocation and light partitioning among dominant and subordinate individuals in Xanthium canadense stands. Annals of Botany, 82: 665-673.
Anten, N.P.R., Miyazawa, K., Hikosaka, K., Nagashima, H. & Hirose, T. (1998) Leaf nitrogen distribution in relation to leaf age and photon flux density in dominant and subordinate plants in dense stands of a dicotyledonous herb. Oecologia, 113: 314-324.
Hikosaka, K., Hanba, Y., Hirose, T. & Terashima, I. (1998) Photosynthetic nitrogen use efficiency in leaves of woody and herbaceous species. Functional Ecology, 12: 896-905.
Hikosaka, K. & Hirose, T. (1998) Leaf and canopy photosynthesis of C3 plants at elevated CO2 in relation to optimal partitioning of nitrogen among photosynthetic components: theoretical prediction. Ecological Modelling, 106: 247-259.
Hikosaka, K., Murakami, A. & Hirose, T. (1998) Temperature acclimation of the photosynthetic apparatus: balancing regeneration and carboxylation of ribulose bisphosphate. In: ed. G. Garab, Photosynthesis: Mechanisms and effects. Vol. V, pp. 3395-3397. Kluwer Academic Press, Dordrecht.
Hirose, T. & Bazzaz, F.A. (1998) Trade-off relationship between light- and nitrogen-use efficiency in canopy photosynthesis. Annals of Botany, 82: 195-202
Shitaka, Y. & Hirose, T. (1998) Effects of shift in flowering time on the reproductive output of Xanthium canadense in a seasonal environment. Oecologia, 114: 361-367.
Sakai, A. & Sakai, S. (1998) A test for the resource recovery hypothesis: tree sprouting using carbohydrates from above-ground parts. Annals of Botany, 82: 213-216.
Sakai, S. (1998) Female-biased Sexual allocation in relation to growth rate of fruit in Erythronium japonicum. Oecologia, 117: 391-395.
Sakai, S. & Harada, Y. (1998) Does the trade-off between growth and reproduction select for female-biased sexual allocation in cosexual plants? Evolution, 52:1204-1207.
Sakai, S., Kikuzawa, K. & Umeki, K. (1998) Evolutionarily stable resource allocation for production of wind-dispersed seeds. Evolutionary Ecology, 12: 477-485.



Anten, N.P.R. (1997) Modelling canopy photosynthesis using parameters determined from simple non-destructive measurements. Ecological Research, 12: 77-88.
Funayama, S., Hikosaka, K. & Yahara, T. (1997) The effects of virus infection and growth irradiance on fitness components and photosynthetic properties of Eupatorium makinoi (Compositae). American Journal of Botany, 84: 823-829.
Hikosaka, K. (1997) Modelling optimal temperature acclimation of the photosynthetic apparatus in C3 plants with respect to nitrogen use. Annals of Botany, 80: 721-730.
Hikosaka, K. & Hirose, T. (1997) Leaf angle as a strategy for light competition: Optimal and evolutionarily stable light-extinction coefficient within a canopy. Ecoscience, 4: 501-507.
Hiratsuka, A. (1997) Dimorphic seeds and population structure of Xanthium occidentale in dam reservoir and river population. Ecolgical Review, 23: 261-271.
Hirose, T., Ackerly, D.D., Traw, M.B., Ramseier, D. & Bazzaz, F.A. (1997) Modeling canopy photosynthesis and optimal leaf area index in response to CO2 elevation in annual plant stands. Ecology, 1
Miura, O., Maekawa, R. & Hiratsuka, A. (1997) Geomorphological and pedological characteristics in relation to habitat of Erythronium japonicum in hills. Ecolgical Review, 23:273-282.
Nagamatsu, D. & Miura, O. (1997) Soil disturbance regime in relation to micro-scale landform and its effects on vegetation structure in a hilly area in Japan. Plant Ecology, 133: 191-200.
Okagami, N., Tanno, N., Kawai, S., Hiratsuka, A., Satou, K. & Terui, K. (1997) Expansion of trait of gibberellin-induced dormancy in the Tertiary relict species of the genus Dioscorea (Dioscoreaceae). Plant Growth Regulation. 22: 137-140.
Sakai, A., Sakai, S., & Akiyama, F. (1997) Do sprouting tree species on erosional sites have large reserves of resources? Annals of Botany, 79: 625-630.
Sakai, S. (1997) An analytical model for flower production in an environment where resource availability is unpredictable. Journal of theoretical Biology, 189: 449-452.
Sakai, S., Sakai, A., & Ishii, H.S. (1997) Patterns of wing size variation in seeds of the lily Cardiocrinum cordatum (Liliaceae). American Journal of Botany 84: 1275-1278.


Anten, N.P.R., Hernandez, R. & Medina, E. (1996) The photosynthetic capacity and leaf nitrogen concentration as related to light regime in shade leaves of a montane tropical forest tree, Tetrorchidium rubrivenium. Functional Ecology, 10: 491-500.
Hikosaka, K. (1996) Effects of leaf age, nitrogen nutrition and photon flux density on the organization of the photosynthetic apparatus in leaves of a vine (Ipomoea tricolor Cav.) grown horizontally to avoid mutual shading of leaves. Planta, 198: 144-150.
Hikosaka, K. & Terashima, I. (1996) Nitrogen partitioning among photosynthetic components and its consequence in sun and shade plants. Functional Ecology, 10: 335-343.
Hirose, T., Ackerly, D.D. & Bazzaz, F.A. (1996a) CO2 elevation and canopy development in stands of herbaceous plants. In: Ch. Korner and F.A. Bazzaz (eds.) Carbon Dioxide, Populations, and Communities. Academic Press, San Diego, USA. pp. 413-430.
Hirose, T., Ackerly, D.D., Traw, M.B. & Bazzaz, F.A. (1996b) Effects of CO2 elevation on canopy development in the stands of two co-occurring annuals. Oecologia, 108: 215-223.
Sakai, S. (1996a) Using phylogenies to explain seed size variation among plants. Journal of Evolutionary Biology, 9: 243-252.
Sakai, S. (1996b) On ovule production in an environmental where pollinator or resource availability is unpredictable. Journal of Theoretical Biology, 183: 317-327.
Sakai, S. (1996c) Theoretical approaches on the growth strategy of plants in unpredictable environments. Plant Morphology, 8: 21-26.
Sakai, S. & Sakai, A. (1996) Why is there variation in mean seed size among plants within single populations? test of the fertilization efficiency hypothesis. American Journal of Botany, 83: 1454-1457.


Anten, N.P.R., Schieving, F. & Werger, M.J.A. (1995) Patterns of light and nitrogen distribution in relation to whole canopy carbon gain in C3 and C4 mono- and dicotyledonous species. Oecologia 101: 504-513.
Anten, N.P.R., Schieving, F., Medina, E., Werger, M.J.A. & Schuffelen, P. (1995) Optimal leaf area indices in C3 and C4 mono- and dicotyledonous species at low and high nitrogen availability. Physiologia Plantarum 95: 541-550.
Hikosaka, K. & Terashima, I. (1995) A model of the acclimation of photosynthesis in the leaves of C3 plants to sun and shade with respect to nitrogen use. Plant, Cell & Environment 18: 605-618.
Hiratsuka, A., Fujitaka, T., Satoh, K. & Akimoto, J. (1995) The role of spathe and seed set of Lysichiton camtschatcense. Ecological Review 24: 21-28.
Hirose, T. & Werger, M.J.A. (1995) Canopy structure and photon flux partitioning among species in a herbaceous plant community. Ecology 76: 466-474.
Sakai, A., Ohsawa, T. & Ohsawa, M. (1995) Adaptive significance of sprouting of Euptelea polyandra, a deciduous tree growing on steep slopes with shallow soil. Journal of Plant Research, 9, 108: 377-386.
Sakai, S. (1995a) Evolutionarily stable growth of a sapling which waits for future gap formation under closed canopy. Evolutionary Ecology 9:444-452.
Sakai, S. (1995b) Evolutionarily stable selfing rates of hermaphroditic plants in competing and delayed selfing modes with allocation to attractive structures. Evolution 49:557-564.
Sakai, S. (1995c) Optimal resource allocation to vegetative and sexual reproduction of a plant growing in a spatially varying environment. Journal of theoretical Biology 175: 271-282.
Sakai, S. (1995d) A model for seed size variation among plants. Evolutionary Ecology 9: 495-527.
Sakai, S. & Sakai, A. (1995) Flower size-dependent variation in seed size: theory and a test. American Naturalist 145:918-934
Terashima, I. & Hikosaka, K. (1995) Comparative ecophysiology of leaf and canopy photosynthesis. Plant, Cell & Environment 18: 1111-1128.
Terashima, I., Ishibashi, M., Ono, K. & Hikosaka, K. (1995) Three resistances to CO2 diffusion: leaf-surface water, intercellular spaces and mesophyll cells. In: Mathis P (ed.) Photosynthesis: From Light to Biosphere Vol V, Kluwer Academic Press, Dordrecht, pp. 537-542.


Hiratsuka, A., Fujitaka, T. & Kikuchi, T. (1994) Size structure and seedling recruitment of Lysichiton camtschatcense population. Ecological Review 23:7-15.
Hirose, T. & Werger, M.J.A. (1994) Photosynthetic capacity and nitrogen partitioning among species in the canopy of a herbaceous plant community. Oecologia 100:203-212.
Kita, I., Takatsuki, S., & Tiba, T. (1994) Histological difference between retrograde corpora lutea of pregnancy and those of estrus in sika deer, Cervus nippon. J. Vet. Med. Sci. 56: 309-334.
Padmalal, U.K.G.K. & Takatsuki, S. (1994) Age-sex differences in the diet of Sika deer on Kinkazan Island, northern Japan. Ecological Research, 9: 251-256.
Takatsuki, S. & Goral, T. (1994) Effects of Sika deer on the regeneration of a Fagus crenata forest on Kinkazan Island, northern Japan. Ecological Research 9: 115-120.
Takatsuki, S., Suzuki, K. & Suzuki, I. (1994) A mass-mortality od Sika deer on Kinkazan Island, northern Japan. Ecological Research 9: 215-223.


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